Skull 5 (D4500) — the most complete early-Pleistocene skull

Cranium D4500 (found 2005) with mandible D2600 (2000) forms the most complete early-Pleistocene human skull. It pairs a small braincase (~546 cm³) with a large, projecting face. Published in Science in 2013 by David Lordkipanidze's team, it underpins the still-debated “single evolving lineage” hypothesis of early Homo. Held at the Georgian National Museum.